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Nitrate supplementation

The alternative NO synthesis pathway has also been well studied with reference to exercise physiology. Nitrate and nitrite modulate mitochondrial respiration through NO synthesis. Two recent studies by the same research group showed that plasma NO metabolites like nitrate and nitrite increased significantly after nitrate treatment. The efficiency of human mitochondria, assessed in vitro by oxygen consumption per molecule of ATP produced (phosphate/oxygen ratio), was significantly enhanced with respect to a placebo group after intake of 0.1 mmol/kg sodium nitrate for 3 days.

Interestingly, studies involving beetroot juice have also reported a significant increase in plasma nitrite levels (regarded as markers of NO) after intake of beetroot juice. Larsen et al. reported it as an alternative metabolic pathway for mitochondrial respiration which might explain the decrease in oxygen demand during exercise after ingestion of nitrate-rich food.

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Conclusions

Nitric oxide is a physiological compound of the human body that dilates blood vessels, stimulates hormone release, regulates neurotransmission and acts as a signaling molecule. Nitric oxide is synthesized by NOS-dependent and -independent pathways. Nitric oxide supplementation improves cardiac health, enhances performance during exercise, improves healing, reduces erectile dysfunction, reduces high blood pressure during pregnancy and improves respiratory response.

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The benefits associated with NO have mostly been obtained in moderately trained individuals. Two prevalent NO supplements are L-arginine and L-citrulline. L-arginine is involved in NO synthesis. L-arginine supplements increase respiratory response and enhance performance during exercise. L-citrulline is a L-arginine precursor that is converted into NO by a reaction catalyzed by NOS. L-citrulline combined with malate and other molecules increase working capacity. Many studies on beetroot juice report a significant increase in plasma nitrite levels, a NO marker, after intake of beetroot juice. Similarly, glycine propionyl-L-carnitine and 2-(nitroxy) ethyl 2-amino-3-methylbutanoate are dietary supplements that act as nitric oxide donors, enhance nitric oxide synthesis and may also stimulate NO synthesis through expression of the eNOS gene.